Glossary of mammalian dental topography
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Many different terms have been proposed for features of the tooth crown in mammals.
The structures within the molars receive different names according to their position and morphology. This nomenclature was developed by Henry Fairfield Osborn in 1907 and is, although with many variations, the one that continues today.[1][2][3]
Suffixes
Tooth structures bear suffixes in order to note the type of structure they are and whether they are present in the upper or lower molars.
- The suffix "-cones /-conids" (upper molar/lower molar) is added to the main cusps: Paracone, Metacone, Protocone and Hypocone on the upper molar, and Paraconid, Metaconid, Protoconid, Hypoconid and Entoconid on the lower molar. This name is used for both bunodont and selenodont molars, that is, as many for "buno" pillar-like cusps as for "selenes" crescent-like cusps.
- The suffix "-conule /-conulid" (upper molar/lower molar) is added to the secondary cusps. For example, Metaconule, Hypoconulid.
- The suffix "-style/-stylid" (upper molar/lower molar) is added to the peripheral cusps that are found in the cornices or cingulus of the tooth. These cusps are traditionally named according to their proximity to the main cusps, although some anatomists prefer to name them according to their position on the tooth.
- The suffix "-loph/-lophid" (upper molar/lower molar) is added to the crests that join cusps together. They include in the name one the cusps involved. For example, the hypolophid is the ridge that unites the hypoconid with the entoconid. These ridges often have secondary ridges: the secondary crest of the ectoloph is called crista; antecrochet to that of the protoloph, and crochet to that of the metaloph.
- The suffix "-crista / -cristid" (upper molar / lower molar) is used for the ridges that come out of the cusps but do not connect them with other cusps. It is also used to name the edges of selenes.
- To the structures in the lingual part of the molar, the prefix "ento-" ("internal") is often added, while those of the lingual part are added the prefix "ecto-" ("external"). the mesial part of the molar is often added the prefix "pre-" (from "previous") while those of the distal part are added the prefix "post-" ("posterior"). The mesial part is that which is towards the incisors.
- The suffix "-flexus / -flexid" (upper molar / lower molar) is used for the open valleys in the occlusal surfaces of the hypsodont teeth. When this valleys are enclosed, they are called fossetes/fossetids (upper molar / lower molar). Sometimes they are used also for the folds of the teeth, although the proper name for the folds is sulcus (pl. sulci).
- The cusp prefixes "para-", "meta-", "proto-", "hypo-", etc., are related to the succession and position of the cusps according to the ancient tritubercular theory of the evolution of molars from Cope and Osborn. Although this theory has lost its validity, they continue to use the names for the description of the molars. The prefix "proto" referred to the original cusp which would be homologous to a single cusp tooth according to Osborn, and would be the first cusp to appear not only in evolution but in development.[1] This was criticized early by studies of embryology where it was shown that the first cusp in the embryonic development of the upper molars was the paracone. Later this was shown to be variable and the first cusp to appear in ontogeny would not be related to the evolution of the tooth.[2]
Major features
-
A diagram of generalized tribosphenic molars with notable features labelled. Upper left molar in pink, lower left molar in blue. Mesial (forwards) is to the left. Teeth are depicted in left lateral view (left side and center of the image) or occlusal view (right side of the image).
The positions of tooth features are described along four directions: mesial (forwards, towards the chin), distal (backwards, towards the jaw joint), lingual (inwards, towards the tongue), and buccal or labial (outwards, towards the cheek).
Upper teeth
Therians (marsupials and placentals) ancestrally have roughly triangular upper molars, with the apex pointing lingually (inwards) and the flat edge positioned labially (outwards). This fundamental three-pointed structure is sometimes called the trigon. Three major cusps are almost always present:
- Page Template:Visible anchor/styles.css has no content.Protocone: A major cusp at the lingual apex of the upper molar. In mammals with tribosphenic teeth, the protocone slides into the talonid basin on the lower teeth in order to grind and crush food. Often the largest cusp in the upper molar, though some insectivorous mammals reduce it substantially.
- Page Template:Visible anchor/styles.css has no content.Paracone: A major cusp outwards from the protocone and in front of the metacone.
- Page Template:Visible anchor/styles.css has no content.Metacone: A major cusp outwards from the protocone and behind the paracone.
Other common features include:
- Page Template:Visible anchor/styles.css has no content.Stylar shelf: A shelf or offset surface along the outer edge of the tooth. Often bears minor cusps (stylar cusps or conules).
- Page Template:Visible anchor/styles.css has no content.Parastyle: A forward extension of the stylar shelf which juts out at the mesiolabial (front outer) corner of the tooth, in front of and outwards from the paracone. May bear one or more minor cusps (including the stylocone, which is observed in some extinct mammals).
- Page Template:Visible anchor/styles.css has no content.Metastyle: A rearward extension of the stylar shelf which juts out at the distolabial (rear outer) corner of the tooth, behind and outwards from the metacone. May bear one or more minor cusps.
- Page Template:Visible anchor/styles.css has no content.Mesostyle: A cusp at the midpoint of the stylar shelf.
- Page Template:Visible anchor/styles.css has no content.Ectoflexus: A notch at the midpoint of the stylar shelf.
- Page Template:Visible anchor/styles.css has no content.Paraconule / Protoconule: A minor cusp on the front edge of the tooth, between the protocone and paracone.
- Page Template:Visible anchor/styles.css has no content.Metaconule: A minor cusp on the rear edge of the tooth, between the protocone and metacone.
- Page Template:Visible anchor/styles.css has no content.Hypocone: A fourth major cusp which only becomes widespread in placentals. It develops at the distolingual (rear inner) corner of the tooth, behind the protocone and inwards from the metacone. Mammals with a large hypocone (such as humans) have upper molars with four cusps set in a quadrate (square-shaped) arrangement rather than a triangular form. Many marsupials and artiodactyls have an enlarged metaconule which serves the same role as the hypocone and is occasionally labelled as a hypocone, despite being derived from a different part of the tooth.[4]
Lower teeth
Therians ancestrally have lower molars which are longer from front-to-back than from side-to-side. Five to six cusps are most common. The trigonid region at the front part of the molar is triangular, with three large cusps:
- Page Template:Visible anchor/styles.css has no content.Protoconid: A major cusp at the labial apex of the trigonid region. Often the largest cusp in the lower molar.
- Page Template:Visible anchor/styles.css has no content.Paraconid: A major cusp on the inner edge of the trigonid region, inwards from the protoconid and in front of the metaconid. Humans and most other primates no longer possess a paraconid in their lower molars, leaving only five cusps.
- Page Template:Visible anchor/styles.css has no content.Metaconid: A major cusp on the inner edge of the trigonid region, inwards from the protoconid and behind the paraconid.
The talonid region at the rear part of the molar has two to three relatively small cusps which define the rear rim of a low basin:
- Page Template:Visible anchor/styles.css has no content.Hypoconid: A major cusp at the outer rear edge of the talonid region. Typically the largest and most consistently present cusp in the talonid, though usually smaller than the trigonid cusps. The hypoconid occludes with the space at the center of the upper molar (between the protocone, paracone, and metacone).
- Page Template:Visible anchor/styles.css has no content.Entoconid: A major cusp which is sometimes present at the inner rear edge of the talonid region.
- Page Template:Visible anchor/styles.css has no content.Hypoconulid: A minor cusp which is sometimes present at the rear edge of the talonid region, between the hypoconid and entoconid.
Other common features include:
- Page Template:Visible anchor/styles.css has no content.Cristid obliqua: A diagonal crest which runs from the hypoconid forwards to the midpoint between the protoconid and metaconid.
Image gallery
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Lophodont teeth with ectoloph, protoloph and metaloph, several small cristae, and an entoflexus and postflexus. -
The bunolophoselenodont dentition of a horse. -
Cricetidae molar teeth nomenclature after Reig (1977)
These are a list of tooth features identified in the rodent family Cricetidae (hamsters, voles, New World mice and rats, etc.). Source:[5]
Upper teeth
| Name | Nomenclature term is used in | Definition | Comments | Image |
|---|---|---|---|---|
| Page Template:Visible anchor/styles.css has no content.Anterolingual conule | Reig (1977)[5] | A conule on the lingual side of an anterocone divided by an anteromedian flexus or fossette | ||
| Page Template:Visible anchor/styles.css has no content.Anteromedian flexus | Reig (1977)[5] | A longitudinal flexus dividing the anterocone into anterolabial and anterolingual conules | ||
| Page Template:Visible anchor/styles.css has no content.Anterolabial conule | Reig (1977)[5] | A conule on the labial side of an anterocone divided by an anteromedian flexus or fossette | ||
| Page Template:Visible anchor/styles.css has no content.Anterocone | Reig (1977)[5] | A cusp at the front of the tooth that may be divided into anterolabial and anterolingual conules | ||
| Page Template:Visible anchor/styles.css has no content.Protostyle | Reig (1977)[5] | A style in front of the protocone, in the protoflexus | ||
| Page Template:Visible anchor/styles.css has no content.Protoflexus | Reig (1977)[5] | A flexus between the protocone and the anterolingual conule | ||
| Page Template:Visible anchor/styles.css has no content.Anterior mure | Reig (1977)[5] | A crest connecting the anterocone to the protocone | ||
| Protocone | Reig (1977)[5] | One of the main cusps, at the anterolingual side | ||
| Page Template:Visible anchor/styles.css has no content.Enterostyle | Reig (1977)[5] | A style between the protocone and the hypocone, in the hypoflexus | ||
| Page Template:Visible anchor/styles.css has no content.Enteroloph | Reig (1977)[5] | A crest connecting the enterostyle to the mesocone | ||
| Page Template:Visible anchor/styles.css has no content.Hypoflexus | Reig (1977)[5] | A flexus between the protocone and the hypocone | ||
| Page Template:Visible anchor/styles.css has no content.Mesocone | Reig (1977)[5] | A conule in the median mure where the mesoloph is attached to it | ||
| Page Template:Visible anchor/styles.css has no content.Median mure | Reig (1977)[5] | A crest connecting the protocone/paracone to the hypocone/metacone | ||
| Hypocone | Reig (1977)[5] | One of the main cusps, at the posterolingual side | ||
| Page Template:Visible anchor/styles.css has no content.Procingulum | Reig (1977)[5] | The front part of the tooth, before the anterior mure | ||
| Page Template:Visible anchor/styles.css has no content.Anteroflexus | Reig (1977)[5] | A flexus between the anteroloph and the anterolabial conule | ||
| Page Template:Visible anchor/styles.css has no content.Anteroloph | Reig (1977)[5] | A crest between the paracone and the anterolabial conule that may be connecting to a parastyle | ||
| Parastyle | Reig (1977)[5] | A style in front of the paracone | ||
| Page Template:Visible anchor/styles.css has no content.Paraflexus | Reig (1977)[5] | A flexus in front of the paracone | ||
| Page Template:Visible anchor/styles.css has no content.Protolophule | Reig (1977)[5] | A small crest in the paraflexus, connected to the protocone | ||
| Page Template:Visible anchor/styles.css has no content.Paraloph | Reig (1977)[5] | A crest attaching the paracone to the protocone or the median mure | ||
| Paracone | Reig (1977)[5] | One of the main cusps, at the anterolabial side | ||
| Page Template:Visible anchor/styles.css has no content.Paralophule | Reig (1977)[5] | A small crest attached to the back side of the paracone | ||
| Page Template:Visible anchor/styles.css has no content.Mesoflexus | Reig (1977)[5] | A flexus between the mesoloph and the paracone | ||
| Mesostyle | Reig (1977)[5] | A style at the labial margin between the paracone and metacone | ||
| Page Template:Visible anchor/styles.css has no content.Mesoloph | Reig (1977)[5] | A crest in front of the metaflexus, connected to the median mure | ||
| Page Template:Visible anchor/styles.css has no content.Metaflexus | Reig (1977)[5] | A flexus in front of the metacone | ||
| Page Template:Visible anchor/styles.css has no content.Metalophule | Reig (1977)[5] | A small crest attached to the front side of the metacone | ||
| Metacone | Reig (1977)[5] | One of the main cusps, at the posterolabial side | ||
| Page Template:Visible anchor/styles.css has no content.Metaloph | Reig (1977)[5] | A crest attaching the paracone to the hypocone | ||
| Page Template:Visible anchor/styles.css has no content.Posteroflexus | Reig (1977)[5] | A flexus between the posteroloph and the metacone | ||
| Page Template:Visible anchor/styles.css has no content.Posterostyle | Reig (1977)[5] | A crest on the posterolabial corner of the molar | ||
| Page Template:Visible anchor/styles.css has no content.Posteroloph | Reig (1977)[5] | A crest at the back of the molar, connected to the hypocone |
Lower teeth
| Name | Nomenclature term is used in | Definition | Comments | Image |
|---|---|---|---|---|
| Page Template:Visible anchor/styles.css has no content.Anterolingual conulid | Reig (1977)[5] | A conulid on the lingual side of an anteroconid divided by an anteromedian flexid or fossettid | ||
| Page Template:Visible anchor/styles.css has no content.Anteromedian flexid | Reig (1977)[5] | A longitudinal flexid dividing the anteroconid into anterolabial and anterolingual conules | ||
| Page Template:Visible anchor/styles.css has no content.Anterolabial conulid | Reig (1977)[5] | A conulid on the labial side of an anteroconid divided by an anteromedian flexid or fossettid | ||
| Page Template:Visible anchor/styles.css has no content.Anteroconid | Reig (1977)[5] | A cusp at the front of the tooth that may be divided into anterolabial and anterolingual conulids | ||
| Page Template:Visible anchor/styles.css has no content.Anterolabial cingulum | Reig (1977)[5] | A crest before the protoconid and protoflexid | ||
| Page Template:Visible anchor/styles.css has no content.Protostylid | Reig (1977)[5] | A stylid in front of the protoconid, in the protoflexid | ||
| Page Template:Visible anchor/styles.css has no content.Protoflexid | Reig (1977)[5] | A flexid between the protoconid and the anterolabial conulid | ||
| Page Template:Visible anchor/styles.css has no content.Anterior murid | Reig (1977)[5] | A crest connecting the anteroconid to the protoconid | ||
| Protoconid | Reig (1977)[5] | One of the main cusps, at the anterolabial side | ||
| Page Template:Visible anchor/styles.css has no content.Ectostylid | Reig (1977)[5] | A stylid between the protoconid and the hypoconid, in the hypoflexid | ||
| Page Template:Visible anchor/styles.css has no content.Ectolophid | Reig (1977)[5] | A crest connecting the ectostylid to the mesoconid | ||
| Page Template:Visible anchor/styles.css has no content.Hypoflexid | Reig (1977)[5] | A flexid between the protoconid and the hypoconid | ||
| Page Template:Visible anchor/styles.css has no content.Mesoconid | Reig (1977)[5] | A conulid in the median murid where the mesolophid is attached to it | ||
| Page Template:Visible anchor/styles.css has no content.Median murid | Reig (1977)[5] | A crest connecting the protoconid/metaconid to the hypoconid/entoconid | ||
| Hypoconid | Reig (1977)[5] | One of the main cusps, at the posterolabial side | ||
| Page Template:Visible anchor/styles.css has no content.Procingulum | Reig (1977)[5] | The front part of the tooth, before the anterior murid | ||
| Page Template:Visible anchor/styles.css has no content.Anteroflexid | Reig (1977)[5] | A flexid between the anterolophid and the anterolingual conulid | ||
| Page Template:Visible anchor/styles.css has no content.Anterolophid | Reig (1977)[5] | A crest between the metaconid and the anterolabial conulid that may be connecting to a metastylid | ||
| Page Template:Visible anchor/styles.css has no content.Metastylid | Reig (1977)[5] | A stylid in front of the metaconid | ||
| Page Template:Visible anchor/styles.css has no content.Metaflexid | Reig (1977)[5] | A flexid in front of the metaconid | ||
| Page Template:Visible anchor/styles.css has no content.Protolophulid | Reig (1977)[5] | A small crest in the mesoflexid, connected to the protoconid | ||
| Page Template:Visible anchor/styles.css has no content.Metalophid | Reig (1977)[5] | A crest attaching the metaconid to the protoconid or the anterior murid | ||
| Metaconid | Reig (1977)[5] | One of the main cusps, at the anterolingual side | ||
| Page Template:Visible anchor/styles.css has no content.Metalophulid | Reig (1977)[5] | A small crest attached to the back side of the metaconid | ||
| Page Template:Visible anchor/styles.css has no content.Mesoflexid | Reig (1977)[5] | A flexid between the mesolophid and the paraconid | ||
| Page Template:Visible anchor/styles.css has no content.Mesostylid | Reig (1977)[5] | A stylid at the labial margin between the metaconid and entoconid | ||
| Page Template:Visible anchor/styles.css has no content.Mesolophid | Reig (1977)[5] | A crest in front of the entoflexid, connected to the median murid | ||
| Page Template:Visible anchor/styles.css has no content.Entoflexid | Reig (1977)[5] | A flexid in front of the entoconid | ||
| Page Template:Visible anchor/styles.css has no content.Entolophulid | Reig (1977)[5] | A small crest attached to the front side of the entoconid | ||
| Entoconid | Reig (1977)[5] | One of the main cusps, at the posterolingual side | ||
| Page Template:Visible anchor/styles.css has no content.Entolophid | Reig (1977)[5] | A crest attaching the entoconid to the hypoconid or median murid | ||
| Page Template:Visible anchor/styles.css has no content.Posteroflexid | Reig (1977)[5] | A flexid between the posterolophid and the metaconid | ||
| Page Template:Visible anchor/styles.css has no content.Hypolophulid | Reig (1977)[5] | A small crest in the posteroflexid attached to the posterolophid | ||
| Page Template:Visible anchor/styles.css has no content.Posterostylid | Reig (1977)[5] | A crest on the posterolingual corner of the molar | ||
| Page Template:Visible anchor/styles.css has no content.Posterolophid | Reig (1977)[5] | A crest at the back of the molar, connected to the hypoconid |
See also
References
Page Template:Reflist/styles.css has no content.
- ^ a b Page Module:Citation/CS1/styles.css has no content.Osborn, Henry Fairfield, 1857–1935. (1907). Evolution of mammalian molar teeth to and from the triangular type: including collected and revised researches on trituberculy, and new sections on the forms and homologies of the molar teeth in the different orders of mammals. Kessinger Publishing. ISBN 978-1-4368-4090-3. OCLC 810943856.
{{cite book}}: ISBN / Date incompatibility (help)CS1 maint: multiple names: authors list (link) CS1 maint: numeric names: authors list (link) - ^ a b Page Module:Citation/CS1/styles.css has no content.MacCord, Katherine (2017). Development, evolution, and teeth: how we came to explain the morphological evolution of the mammalian dentition. Arizona State University.
- ^ Page Module:Citation/CS1/styles.css has no content.Szalay, Frederick S. (1969). "Mixodectidae, Microsyopidae, and the insectivore-primate transition". Bulletin of the American Museum of Natural History. 140 4: 193–330. hdl:2246/1130.
- ^ Page Module:Citation/CS1/styles.css has no content.Hunter, J P; Jernvall, J (7 November 1995). "The hypocone as a key innovation in mammalian evolution". Proceedings of the National Academy of Sciences. 92 (23): 10718–10722. Bibcode:1995PNAS...9210718H. doi:10.1073/pnas.92.23.10718. ISSN 0027-8424. PMC 40683. PMID 7479871.
- ^ a b c d e f g h i j k l m n o p q r s t u v w x y z aa ab ac ad ae af ag ah ai aj ak al am an ao ap aq ar as at au av aw ax ay az ba bb bc bd be bf bg bh bi bj bk bl bm bn bo bp bq Page Module:Citation/CS1/styles.css has no content.Reig, Osvaldo A. (2009). "A proposed unified nomenclature for the enamelled components of the molar teeth of the Cricetidae (Rodentia)". Journal of Zoology. 181 (2): 227–241. doi:10.1111/j.1469-7998.1977.tb03238.x. ISSN 0952-8369.
