Homalozoa
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Homalozoa is an obsolete extinct subphylum of Paleozoic era echinoderms, prehistoric marine invertebrates. They are also referred to as carpoids.[1]
Description
The Homalozoa lacked the typical pentamer body form of other echinoderms, but all were sessile animals. Instead all Homalozoans were markedly asymmetric, and were extremely variable in forms.
The body (theca) was covered with calcite plates with a number of openings. Their form is in some cases so unusual that it is unclear which openings are to be considered as mouth and anus. Many of them were stalked, similar to sea lilies (crinoids), but often their bodies were bent over, so that the mouth and anus projected forwards rather than upwards. Some forms, especially stylophorans, rested flat on the sea floor.[2]
In some forms the single ray (brachiole or aulacophore) possessed an ambulacral groove.[3]
It has been claimed that some forms possessed gills and gill slits.[4]
The first homalozoans found in the fossil record are in lower Middle Cambrian rocks of northern Utah.[5]
Taxonomy
Homalozoans were traditionally considered to be stem-group echinoderms,[6] but had also been considered to lie in the stem lineage of the chordates (calcichordates). However, it is now generally accepted that homalozoans were echinoderms because their calcite skeleton was composed of the typical stereom crystalline structure.[7]
They include the unusual stylophorans (mitrates and cornutes), Homoiostelea (solutes), the Homostelea (cinctans), and the Ctenocystoidea (ctenoid-bearing homalozoans).[8] They have recently been recognised as a polyphyletic group. The stylophorans are now classified as a clade of the Crinozoa, whereas the other three are classified as clades of the Blastozoa.[8]
Solutes
Unlike many other types of echinoderm, solute homalozoans lack radial symmetry (such as the five limbs of a starfish).[9] [10] Solutes are the sole order of the class Homoiostelea.
Solute fossils have an irregularly shaped flattened body covered in calcite plates, and are up to about 10 cm long. The body has two appendages, interpreted as a "feeding arm" at one end, bearing tube feet at its end, and a "stele" at the other, which may have been used by the animal to propel itself along the sea floor.[11]
See also
References
Page Template:Reflist/styles.css has no content.
- ^ Page Module:Citation/CS1/styles.css has no content.Imran Rahman (January–February 2009). "Making sense of carpoids". Geology Today. 25 (1): 34–38. Bibcode:2009GeolT..25...34R. doi:10.1111/j.1365-2451.2009.00703.x.
- ^ Page Module:Citation/CS1/styles.css has no content.Lefebvre, Bertrand (2003). "Functional Morphology of Stylophoran Echinoderms". Palaeontology. 46 (3): 511–555. Bibcode:2003Palgy..46..511L. doi:10.1111/1475-4983.00309.
- ^ Page Module:Citation/CS1/styles.css has no content.Barnes, Robert D. (1982). Invertebrate Zoology. Philadelphia, PA: Holt-Saunders International. p. 1011. ISBN 978-0-03-056747-6.
- ^ Page Module:Citation/CS1/styles.css has no content.Dominguez, Patrício; Jacobson, Antone G.; Jefferies, Richard P. S. (2002). "Paired gill slits in a fossil with a calcite skeleton". Nature. 417 (6891): 841–844. Bibcode:2002Natur.417..841D. doi:10.1038/nature00805. PMID 12075349.
- ^ Page Module:Citation/CS1/styles.css has no content.Robison, R. A.; Sprinkle, J. (December 19, 1969). "Ctenocystoidea: new class of primitive echinoderms". Science. 166 (3912): 1512–1514. doi:10.1126/science.166.3912.1512. PMID 17742852.
- ^ Page Module:Citation/CS1/styles.css has no content.James W. Valentine (2004). On the origin of phyla. University Chicago Press. 608 pp. Paperback. ISBN 978-0-226-84548-7. - pages 401-404
- ^ Page Module:Citation/CS1/styles.css has no content.UCMP Berkeley, edu. "Echinodermata: Morphology". University of California Museum of Paleontology. Retrieved 21 March 2011.
- ^ a b Page Module:Citation/CS1/styles.css has no content.David, Bruno; Lefebvre, Bertrand; Mooi, Rich; Parsley, Ronald (2000). "Are homalozoans echinoderms? An answer from the extraxial-axial theory". Paleobiology. 26 (4): 529–555. Bibcode:2000Pbio...26..529D. doi:10.1666/0094-8373(2000)026<0529:AHEAAF>2.0.CO;2.
- ^ A. B. Smith Deuterostome phylogeny and the interpretation of problematic fossil echinoderms, page 543-544 in Thomas Heinzeller, James H. Nebelsick Echinoderms: München, CRC Press, 2004 Template:ISBN,
- ^ Page Module:Citation/CS1/styles.css has no content.Smith, A. B. (2005). "The pre-radial history of echinoderms". Geological Journal. 40 (3): 255–280. Bibcode:2005GeolJ..40..255S. doi:10.1002/gj.1018.
- ^ Henry Gee Before the backbone: views on the origin of the vertebrates, Springer, 1996 Template:ISBN page 204
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