Heppia
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Heppia is a genus of olive, brownish, grey, or blackish squamulose, crustose, or peltate like lichens.[2] Heppia was once the type genus of the family Heppiaceae, but that family was folded into synonymy with Lichinaceae.[3]
In a multilocus phylogeny and re-classification of the class Lichinomycetes published in 2024, María Prieto, Mats Wedin and Matthias Schultz placed Heppia in the family Porocyphaceae as part of an emended, broader circumscription of that family; earlier treatments that folded Heppiaceae into Lichinaceae are therefore superseded by the new arrangement. Species of Heppia are cyanolichens (usually with filamentous cyanobacteria) that favour open, well-lit habitats on soil or rock, including biological soil crusts in dry regions; they characteristically produce small Template:Lichengloss or peltate to crustose thalli and develop predominantly pycnoascocarpous sexual structures, consistent with the family diagnosis.[4]
The genus was circumscribed by Carl Wilhelm von Nägeli and Abramo Bartolommeo Massalongo in 1854.[5] The genus name Heppiella honours Johann Adam Philipp Hepp (1797–1867), a German physician and lichenologist.[6]
Description
Heppia species grow on rock or soil in arid sites around the world, in habitats similar to those favoured by Peltula, which is similar but has a different cyanobacterium as the Template:Lichengloss.[2] It lacks a medulla that is separate from the photobiont layer.[2] It is a cyanolichen with the photobiont cyanobacterium being Syctonema (or Syctonema-like).[2] The lower surface is paler than upper surface, and has numerous rhizoidal hyphae attaching it to the substrate.[2] The fruiting structures (ascomata) are apothecias immersed in the thallus with red to red-brown urn shaped (Template:Lichengloss) to flat or slightly convex discs.[2] An exciple may or may not be present.[2]
Species
- Heppia adglutinata A.Massal. (1854)
- Heppia arenacea M.Schultz (2005)
- Heppia conchiloba Zahlbr. (1902)
- Heppia despreauxii (Mont.) Tuck. (1872)
- Heppia lutosa (Ach.) Nyl. (1869)
References
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- ^ Page Module:Citation/CS1/styles.css has no content."Synonymy: Heppia Nägeli ex A. Massal". Species Fungorum. CAB International. Retrieved 31 October 2014.
- ^ a b c d e f g Lichen Flora of the Greater Sonoran Desert Region. Vol 1, Nash, T.H., Ryan, B.D., Gries, C., Bugartz, F., (eds.) 2001, [1]
- ^ Page Module:Citation/CS1/styles.css has no content.Schultz, Matthias; Büdel, Burkhard (2003). "On the systematic position of the lichen genus Heppia". The Lichenologist. 35 (2): 151–156. Bibcode:2003ThLic..35..151S. doi:10.1016/s0024-2829(03)00019-7.
- ^ Page Module:Citation/CS1/styles.css has no content.Prieto, M.; Wedin, M.; Schultz, M. (2024). "Phylogeny, evolution and a re-classification of the Lichinomycetes". Studies in Mycology. 109: 595–655. doi:10.3114/sim.2024.109.09. PMC 11663425. PMID 39717657.
- ^ Page Module:Citation/CS1/styles.css has no content.Massalongo, A. (1854). Geneacaena lichenum noviter proposita ac descripta [A newly proposed and described lineage of lichens] (in Latina). Verona: Typic Amanzinianis. pp. 7–8.
- ^ Page Module:Citation/CS1/styles.css has no content.Hertel, Hannes (2012). Gattungseponyme bei Flechten und Lichenicolen Pilzen [Genus eponyms among lichens and lichenicolous fungi]. Bibliotheca Lichenologica (in Deutsch). Vol. 107. Stuttgart: J. Cramer. pp. 50–51. ISBN 978-3-443-58086-5.
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