Pacullidae
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Pacullidae is a family of araneomorph spiders first described by Eugène Simon in 1894.[1] It was merged into Tetrablemmidae in 1958,[2][3] then raised back to family status after a large phylogenetic study in 2017.[4]
Description
The family Pacullidae contains three-clawed spiders with six eyes, lacking a cribellum. They resemble spiders from the family Tetrablemmidae in some respects but are much larger, always exceeding 5 mm long, have a very wrinkled (rugose) cuticle, and females do not have large membranous receptacles.[4]
Phylogeny
Pacullidae falls within the Synspermiata clade, a clade of former haplogyne spiders with "synsperm" – encapsulated groups of 2–4 fused sperm cells. Within this clade, it groups with four other families, including Tetrablemmidae, but is distinct from the latter, being most closely related to Diguetidae. Together with Pholcidae, these four families are placed in the "lost trachea clade", a group of families that have lost their posterior respiratory system.[4] Script error: No such module "Clade".
Genera
As of January 2026[update], this family includes four genera and 38 species:[5]
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Extinct
- †Furcembolus Wunderlich 2008 9 spp, Burmese amber, Myanmar, Cenomanian
References
- ^ Page Module:Citation/CS1/styles.css has no content.Simon, E. (1894). Histoire naturelle des araignées.
- ^ Page Module:Citation/CS1/styles.css has no content.Bonnet, P. (1958). "Bibliographia araneorum". Douladoure, Toulouse. 2 (4): 3297.
- ^ Page Module:Citation/CS1/styles.css has no content.Brignoli, P. M. (1973). "Ragni della Melanesia, I. Un nuovo Tetrablemma di Guadalcanal (Isole Salomone) (Araneae Tetrablemmidae)". Memorie della Società Entomologica Italiana, Genova. 72: 86.
- ^ a b c Page Module:Citation/CS1/styles.css has no content.Wheeler, Ward C.; et al. (2016), "The spider tree of life: phylogeny of Araneae based on target-gene analyses from an extensive taxon sampling", Cladistics, 33 (6): 574–616, doi:10.1111/cla.12182, PMID 34724759, S2CID 35535038
- ^ Page Module:Citation/CS1/styles.css has no content."Family Pacullidae Simon, 1894". World Spider Catalog. doi:10.24436/2. Retrieved 2026-01-17.
External links
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